王文采

(中國(guó)科學(xué)院植物研究所系統(tǒng)與進(jìn)化植物學(xué)國(guó)家重點(diǎn)實(shí)驗(yàn)室，北京 100093)

圖1 A～F.隆子翠雀花：A.莖下部；B.復(fù)總狀花序；C.花瓣頂端；D.退化雄蕊；E.雄蕊；F.雌蕊群 G～K.黃花翠雀花：G.植株全形；H.花瓣頂端；I.退化雄蕊；J.雄蕊；K.雌蕊群Fig.1 A-F.Delphinium longziense: A.Lower part of stem; B.Compound raceme; C.Petal apex; D.Staminode; E.Stamen; F.Gynoecium(from holotype) G-K. D.xanthanthum: G.Habit; H.Petal apex; I.Staminode; J.Stamen; K.Gynoecium(from holotype)

中國(guó)科學(xué)院植物研究所標(biāo)本館(PE)派隊(duì)于2017年8～9月在西藏東南部隆子縣一帶進(jìn)行了深入采集，采集到大量植物標(biāo)本，作者在鑒定其中毛茛科翠雀花屬(Delphinium)[1～6]植物標(biāo)本的過(guò)程中發(fā)現(xiàn)了該屬以下七新種。

隆子翠雀花 圖1：A～F

DelphiniumlongzienseW.T.Wang, sp. nov. Fig.1:A-F.

(Sect.ElatopsisHuth)

In Sinae Sect.ElatopseHuth, species circa sexaginta sepala caerulea vel purpurea habent, et eae tres,D.pseudocandelabrumW.T.Wang,D.xanthanthumW.T.Wang etD.longzienseW.T.Wang, sepalis flavis praeditae sunt. Ob sepala flava, petala et staminoida nigra species tres hae inter sese fortasse affines sunt. In eis,D.longziensefloribus in racemos compositos multi-floros dispositis aD.pseudocandelabroetD.xanthantho, quorum inflorescentiae corymbosae 2-3-florae sunt, praeclare differt.

Perennial herbs. Stems 5-11 cm tall, spreading or slightly retrorsely pubescent, simple, 2-leaved. Basal leaves 1-2, long petiolate; blades papery, reniform, 2-3.4 cm×4.5-5.5 cm, at base broadly cordate, on both surfaces on nerves appressed-pubescent, 3-lobed, central lobe obovate-quadrate, 1.2-1.8 cm broad, 3-lobulate, with lobules broad-ovate entire or 1-denticulate and at apex rounded or obtuse, lateral lobes larger, subflabellate, 2.8-3.5 cm broad, unequally 3-lobed; petioles 7-8 cm long, spreading-pubescent. Cauline leaves similar to basal ones, but slightly smaller. Compound racemes terminal, 30-40 cm long; terminal raceme 10-12 cm long, 5-6-flowered and lateral ones 7-13 cm long, 2-4-flowered; rachis with pedicels densely spreading-puberulous; lower bracts foliaceous, upper ones obovate or spatulate, 0.8-1.2 cm long, undividied; pedicels 2-4.5 cm long, near apex 2-bracteolate; bracteoles narrow-oblong or narrow-elliptic, 3-6 mm×2-2.5 mm. Flower: Sepals yellow, inside sparsely and outside slightly densely yellow-sericeous, upper sepal broad-ovate, 10-12 mm×8 mm, spur straight or slightly decurved, subulate, 12-14 mm long, at base 3 mm across, lateral sepals obliquely obovate, 11 mm×9 mm, lower sepals similar to lateral ones, but longer, 15 mm×8 mm. Petals 2, black, at apex 2-lobed, lobes narrow-triangular, 1 mm long, with few long hairs. Staminodes 2, black; limbs narrow-oblong, 6.2 mm×4 mm, 2-fid nearly to the middle of limb, adaxially long yellow-barbate, lobes long ciliate; claws 7 mm long, not appendiculate. Stamens numerous, glabrous; filaments 5 mm long, below lanceolate-linear, above filiform; anthers brownish, oblong, 1.8-2 mm×0.8-1 mm. Carpels 3; ovaries 4.2-4.5 mm×0.8 mm, only on ventral sutures pubescent; styles 1.6-1.8 mm long, glabrous.

西藏(Xizang)：隆子縣，扎日鄉(xiāng)，從桑巴村至曲桑村途中(Lhünzê Xian, Zhari Xiang, on the way from Sangba Cun to Qusang Cun)，alt. 3 460 m，林邊，草本植物，花淡綠黃色(at forest margin;herbs with flowers greenish-yellow)，2017-09-05，PE西藏考察隊(duì)(PE Xizang Exped.)5921(02293621)(holotype,PE)。

我國(guó)翠雀花屬的密花翠雀花組(Sect.ElatopsisHuth)有約60種具藍(lán)色或紫色萼片，有3種，石灘翠雀花(D.pseudocandelabrumW.T.Wang)，黃花翠雀花(D.xanthanthumW.T.Wang)和隆子翠雀花(D.longzienseW.T.Wang)，則具黃色萼片。此三種具有以下共同特征：萼片黃色，花瓣和退化雄蕊黑色，這說(shuō)明三種間具有親緣關(guān)系，其中，隆子翠雀花具多花的復(fù)總狀花序，而與其他二種區(qū)別。在其他二種，花序?yàn)榫?或3花的傘房花序(見(jiàn)下)。

黃花翠雀花 圖1：G～K

DelphiniumxanthanthumW.T.Wang, sp. nov. Fig.1:G-K.

(Sect.ElatopsisHuth)

Ob sepala flava, petala et staminodia nigra, inflorescentias corymbosas 2-3-floras species nova haec est affinisD.pseudocandelabroW.T.Wang, a quo foliis minoribus, calcare sepalino sepalis leviter breviore vel eis subaequilongo, petalis apice 2-lobatis carpellis glabris distinguitur. InD.pseudocandelabro, folia majora cum laminis 3-4 cm latis, calcar sepalinum sepalis multo brevius, petala apice integra, et carpellorum ovaria pubescentia sunt.

Perennial herb. Stem 10 cm tall, simple, densely spreading-pubescent, ca. 2-leaved. Basal leaf solitary, long petiolate; blade papery, reniform, 1.2 cm×2.5 cm, at base cordate, on both surfaces sparsely appressed-pubescent, 3-lobed, lobes among themselves contiguous, central lobe small, broadly obtrapeziform, 8 mm broad, at apex 3-dentate, lateral lobes flabellate, 2 cm broad, unequally 3-lobulate, lobules at apex 1-3-rotund-dentate; petiole 2 cm long, densely retrorse-pubescent. Cauline leaves shortly petiolate; blades pentagonal, 0.8-1 cm×1.4-1.7 cm, 3-lobed nearly to the middle of blade, central lobe broad-ovate, lateral lobes obliquely broad-obovate, unequally 2-lobed, with secondery lobes ovate and entire; petioles 0.6-1 cm long. Corymb terminal, 3.5 cm long, 2-flowered; basal bract 5.5 mm long, 3-lobed, upper bract narrow-oblanceolate, 10 mm long, undivided; pedicels 3.5 cm long, densely spreading-puberulous, above the middle or near-flower 2-bracteolate; bracteoles linear, 4.5-6 mm×1-2 mm. Flower: Sepals yellow, inside sparsely, outside slightly densely yellowish-sericeous, upper sepal suborbicular, 12 mm×14 mm, spur thickly subulate, 13-14 mm long, at base 3-4 mm across, lateral sepals broad-obovate, 12×15 , lower sepals obovate 13 mm×11 mm. Petals 2, black, glabrous, apex 2-lobed, lobes linear 2.2-4.5 mm long. Staminodia 2, black; limbs oblong, 10 mm×4.6 mm, 2-fid(lobes lanceolate-linear, 5-6 mm×1.5-1.8 mm), adaxially below lobes and on lobes long yellow-barbate; claws 7 mm long, at base 1-appendiculate, Stamens numerous, glabrous, filaments lanceolate-linear, 5.5-6 mm long; anthers dark-blue, oblong, 1.8-2 mm×0.8-1 mm. Carpels 3, glabrous; ovaries greenish-white, at apex dark-blue, 4 mm×0.8 mm; styles white, 1 mm long.

西藏(Xizang)：隆子縣，扎日鄉(xiāng)，納拉山(Lhünzê Xian,Zhari Xiang,Mt. Nala)，alt. 3 400 m，山坡上(on slopes)，2017-09-03，PE西藏考察隊(duì)(PE Xizang Exped.)5813(02293611)(holotype,PE)。

本種的萼片黃色，花瓣和退化雄蕊黑色，花組成具2花的傘房花序，因此與也具上述特征的石灘翠雀花(D.pseudocandelabrumW.T.Wang)相近緣，與后者的區(qū)別在于本種的葉較小，葉片寬1.4～2.5 cm，萼距比萼片稍短或近等長(zhǎng)，花瓣頂端2淺裂，心皮無(wú)毛。在石灘翠雀花，葉較大，葉片寬3～4 cm，萼距比萼短的多，花瓣頂端全緣，心皮的子房被柔毛。(王文采，1983；Wang & Warnock,2001)

綠房翠雀花 圖2：A～E

DelphiniumviridiovariumW.T.Wang, sp. nov. Fig.2:A-E.

(Sect.ElatopsisHuth)

Ob folia 3-lobata, calcar sepalinum sepalis subaequilongum, petala et staminodia nigra, carpella glabra species nova haec est affinisD.langxianensiW.T.Wang, a quo foliis majoribus, floribus in corymbos 3-floros dispositis, pedicellis supra medium 2-bracteolatis differt. InD.langxianensi, folia minora cum laminis 1-2 cm longis et 1.5-3.6 cm latis, flores in racemos 7-8-floros dispositi, et pedicelli ad apicem 2-bracteolati sunt.

Perennial herb. Stem ca. 20 cm tall, simple, with petioles and pedicels spreading-puberulous, 3-leaved. Basal leaf solitary, long petiolate; blade papery, reniform, ca. 3.8 cm×7 cm, at base cordate, adaxially on nerves appressed-puberulous, abaxially sparsely puberulous, 3-lobed nearly to the middle of blade, central lobe obtrapeziform-obovate, 1.5 cm broad, 3-lobulate with lobules broad-ovate, 1-2-denticulate or entire, lateral lobes subflabellate, 3.5-4 cm broad, unequally 2-lobed, secondary lobes shortly obtrapeziform or broad-obovate, unequally 3-lobulate; petiole 13 cm long, at base shortly vaginate. Cauline leaves similar to basal one, but with smaller blades and shorter petioles. Corymb terminal, ca. 8 cm long, 3-flowered; bracts lanceolate or narrow-lanceolate, 1.2-1.8 cm long; pedicels 3.5-8 cm long, above the middle 2-bracteolate; bracteoles narrow-lanceolate or lanceolate, 5-10 mm long, puberulous. Flower: Sepals blue-purple, inside glabrous, outside sparsely puberulous, upper sepal broad-ovate, 13 mm×10 mm, spur subulate, 13-14 mm long, slightly decurved, at base 4-5 mm across, lateral sepals orbicular-obovate, 11 mm×15 mm, at apex shortly acuminate and ciliate, lower sepals obovate, 18 mm×11 mm. Petals 2, black, glabrous, apex 2-lobed. Staminodes 2, black; limbs ovate-oblong, 7 mm×5 mm, 2-lobed(lobes 3-4 mm long), adaxially whitish-barbate; claws 7 mm long, at base 1-appendiculate. Stamens numerous, glabrous; filaments 6-7.5 mm long, below lanceolate-linear, above filiform; anthers dark-blue, oblong, 1-1.3 mm×0.7-1 mm. Carpels 3, glabrous; ovaries green, 4.8 mm×0.8 mm; styles white, 2-2.4 mm long.

西藏(Xizang)：洛扎縣，拉康鎮(zhèn)，卡久寺附近(Lhozhag Xian,Lakang Zhen,near Kajiu Temple)，alt. 3 934 m，山坡路邊，草本植物(near roadsides on slopes, herbs)，2017-09-14，PE西藏考察隊(duì)(PE Xizang Exped.)6634(02293616)(holotype,PE)。

本種的葉3淺裂，萼距與萼片近等長(zhǎng)，花瓣和退化雄蕊黑色，心皮無(wú)毛，因此與也具上述特征的朗縣翠雀花(D.langxianenseW.T.Wang)相近緣，與后者的區(qū)別在于本種的葉較大，葉片長(zhǎng)達(dá)3.8 cm，寬達(dá)7 cm，花組成具3花的傘房花序，花梗在中部之上具2小苞片。在朗縣翠雀花，葉較小，葉片長(zhǎng)1～2 cm，寬1.5～3.6 cm，花組成具7～8花的總狀花序，花梗在頂端具2小苞片(王文采，2014)。

二距翠雀花 圖2：F～J

DelphiniumdicentrumW.T.Wang, sp. nov. Fig.2:F-J.

(Sect.ElatopsisHuth)

Ob foliorum laminam lobatam et lobos inter sese contiguous, petala et staminodia nigra, pedicellos prope flores 2-bracteolatos species nova haec est similisD.langxianensiW.T.Wang, a quo folii basalis lamina 2-nervi et 2-lobata, sepalo infero uno basi breviter calcarato valde differ. InD.langxianensi, folii lamina 3-nervis et 3-lobata, et sepala infera haud calcarata sunt.

圖2 A～E.綠房翠雀花：A.基生葉；B.莖頂部和傘房花序；C.退化雄蕊；D.雄蕊；E.雌蕊群 F～J.二距翠雀花：F.根狀莖，莖基部和基生葉；G.總狀花序；H.退化雄蕊；I.雄蕊；J.雌蕊群Fig.2 A-E. Delphinium viridiovarium: A.Basal leaf; B.Apical part of stem and corymb; C.Staminode; D.Stamen; E.Gynoecium(from holotype) F-J. D.dicentrum: F.Rhizome,basal part or stem and basal leaf; G.Raceme; H.Staminode; I.Stamen; J.Gynoecium(from holotype)

Perennial herb. Stem ca. 18 cm tall, simple, slightly retrorsely hirtellous, 2-leaved. Basal leaf solitary, long petiolate; blade papery, reniform, 2 cm×4.6 cm, at base cordate, on both surfaces very sparsely hirtellous and below on margins ciliate, 2-lobed, with 2 primary nerves(each primary nerve above base 3-branched), lobes 3-4-lobulate, lobules among themselves contiguous, shortly obtrapeziform or broad-ovate, at apex rounded, entire or 1-rotund-denticulate; petiole 11 cm long, retrorse-hirtellous, at base narrowly vaginate. Cauline leaves not seen. Raceme terminal, ca. 20 cm long, 3-flowered; rachis with pedicels retrorse-hirtellous; bracts shortly petiolate, blades broad-triangular, 2.5-3.5 cm×3-6.5 cm, at base subtruncate, 3-parted, central lobe obtrapeziform, at apex 3-dentate, lateral lobes larger, broad-obovate, unequally 2-lobed; pedicels 0.5-11 cm long, near flower 2-bracteolate; bracteoles linear, 10-12(-16)×1.5-2(-3) mm. Flower: Sepals purplish, inside glabrous, outside appressed-pubescent, upper sepal broad-ovate, 20 mm×15 mm, spur upwards spreading, rarely lecurved, thickly subulate, 13-15 mm long, at base 3-5 mm across, lateral sepals broad-elliptic, 15 mm×10 mm, lower sepals similar to lateral ones, but larger, 20 mm×9-12 mm, and the left lower sepal at base with a short tubular spur 4 mm long and 1.8 mm in diam. Petals 2, black, glabrous, apex 2-lobed. Staminodes 2, black; limbs oblong, 10 mm×4 mm, 2-lobed to the middle, adaxially below lobes and on lower parts of lobes long white-barbate; claws 7 mm long, at base not appendiculate. Stamens numerous, glabrous; filaments 7.5-8 mm long, below lanceolate-linear, above filiform; anthers dark, oblong, 1.4 mm×0.6-0.8 mm. Carpels 3; ovaries 6-8 mm×0.8-1.8 mm, pubescent or subglabrous; styles 2.5-3 mm long, glabrous.

西藏(Xizang)：隆子縣，從玉麥鄉(xiāng)至曲桑村埡口(Lhünzê Xian,on the way from Yumai Xiang to Qusang Cun)，alt. 4 398 m，高山草地上，花淡紫色(on alpine meadow, fls. light purple)，2017-09-07，PE西藏考察隊(duì)(PE Xizang Exped.)6112(02293612)(holotype,PE)。

本種葉的葉片淺裂，裂片互相鄰接，花瓣和退化雄蕊黑色，花梗在頂端近花處具2小苞片，因此與也具上述特征的朗縣翠雀花(D.langxianenseW.T.Wang)相似，但本種有兩個(gè)十分獨(dú)特的特征，即其葉片具2條一級(jí)脈，每條一級(jí)脈在基脈之上具3條分枝，葉片2淺裂，再就是花的左側(cè)下萼片在基部具1短筒狀距。在朗縣翠雀花以及翠雀花屬其他種，葉片具1條中脈和2條側(cè)生基出脈，葉片分裂成3個(gè)一級(jí)裂片，花只有上萼片具1條距，其他的2側(cè)萼片和2下萼片均無(wú)距。

扎囊翠雀花 圖3：A～E

DelphiniumzhanangenseW.T.Wang, sp. nov. Fig.3:A-E.

(Sect.ElatopsisHuth)

Species nova haec est affinisD.hillcoatiaeMunz, a quo caulibus racemorum rhachidibus et pedicellis haud flavor-glanduloso-puberulis, sepalis apice longe acuminates recedit. InD.hillcoatiae, caules racemorum rhachides et pedicelli patule flavor-glanduloso-puberuli et appresse albo-puberuli, sepala apice acuta sunt.

Perennial herbs. Stems 32-80 cm tall, simple or with ca. 2 short vegetative branthes, densely puberulous, 6-10-leaved. Basal leaves not seen. Cauline leaves long petiolate; blades papery, reniform or pentagonal, 3-5.2 cm×4.5-8 cm, at base cordate, on both surfaces sparsely puberulous or subglabrous, 3-fid to the middle of blade or 3-parted, central lobe obtrapeziform, rhombic or broad-rhombic, 1.6-2 cm broad, at apex subrounded or acute, 3-dentate or 3-lobed, secondary lobes 1-4-denticulate, lateral lobes larger, flabellate, unequally 2-lobed, upper secondary lobe similar to primary central lobe, lower secondary lobe once again unequally 2-lobed; petioles 1-7.5 cm long. Racemes terminal, 20-50 cm long, 11-22-flowered; rachis with pedicels densely spreading-puberulous; lower bracts foliaceous, upper ones lanceolate or linear, 0.8-1.5 cm long, undivided; pedicels 1-6 cm long, above the middle to near flower 2-bracteolate; bracteoles narrow-oblong or narrow-spatulate, 6-15 mm×1-2 mm, longitudinally 1-nerved. Flower: Sepals 5, upper sepal purplish, narrow-ovate, 14-18 mm×5-7 mm, at apex long acuminate, inside glabrous, outside sparsely puberulous or subglabrous, on margin ciliolate, spur subulate, 15-18 mm long, at base 4-5 mm across, lateral and lower sepals white, tinged with blue, elliptic-ovate, 14-16 mm×6-8 mm, at apex long acuminate, subglabrous, longtitudinally 3-blue-nerved. Petals 2, black, glabrous, apex 2-lobulate. Staminodes 2, black; limbs ovate, 6 mm×4 mm, at apex 2-lobulate, adaxially below lobules densely yellow-barbate; claws 6 mm long, at base 1-appendiculate. Stamens numerous, glabrous; filaments 5.5-7 mm long, lanceolate-linear, near apex filiform; anthers dark, oblong, 1.2-1.8 mm×0.6-1 mm. Carpels 3; ovaries 4 mm×0.8 mm, puberulous; styles 2.2-2.5 mm long, glabrous.

西藏(Xizang)：扎囊縣，阿扎鄉(xiāng)(Zhanang Xian,Azha Xiang)，alt. 3 788 m，林中，草本植物，花淡紫色(in forest,herbs,fls. light purple)，2017-08-25，PE西藏考察隊(duì)(PE Xizang Exped.)5088(02293615)(holotype,PE)。

本種與毛茛葉翠雀花(D.hillcoatiaeMunz)在親緣關(guān)系上相近，與后者的區(qū)別在于本種的莖、總狀花序軸和花梗均無(wú)黃色短腺毛，以及萼片頂端長(zhǎng)漸尖。在毛茛葉翠雀花，莖、總狀花序軸和花梗均被開(kāi)展的黃色短腺毛和貼伏的白色短柔毛，萼片頂端急尖。(王文采，1979；Wang & Warnock,2001)

寬片翠雀花 圖3：F～J

DelphiniumlatilimbumW.T.Wang, sp. nov. Fig.3:F-J.

(Sect.DelphinastrumDC.)

Species nova haec est affinisD.hirticauliFranch., a quo caulibus glabris vel sparse pubescentibus, racemis 2-4-floris, pedicellis prope apicem dense puberulis, staminodiorum limbis orbiculari-obovatis, ovariis dense pubescentibus distat. InD.hirticauli, caules inferne hirsutis, racemi 5-10-flori, pedicelli glabri, staminodiorum limbi ovati, ovaria glabra vel subglabra sunt.

Perennial herbs. Stems 6-20 cm tall, simple or long 1-3-branched, glabrous or with very sparse hairs. Basal leaf usually solitary, long petiolate; blade papery, reniform, 3.5-5 cm×6.5-10 cm, at base cordate, on both surfaces with very sparse and very short appressed hairs, 3-parted, central lobe cuneate-rhombic or cuneate, 2-4 cm broad, 3-lobed, with secondary lobes lanceolate-oblong, undivided or 2-3-lobulate, at apex often mucronate, lateral lobes flabellate, 3-6.5 cm broad, unequally 2-fid; petiole slender, 8-19 cm long, glabrous, at base narrowly vaginate. Cauline leaves smilar to basal one, but smaller. Racemes terminal, 3-20 cm long, 2-4-flowered; rachis glabrous; basal bract similar to cauline leaves, petiolate, sometimes 3-sect, with linear ultimate lobes, upper bracts narrow-lanceolate or linear, 1-3 cm long; pedicels 2-14.5 cm long, near flower densely spreading-puberulous, above the middle to near flower 2-bracteolate; bracteoles linear or subulate, 7-15 mm×0.5-2 mm, puberulous. Flower: Sepals blue, inside glabrous, only in lower sepals below with a few short hairs, outside appressed-puberulous, upper sepal broad-ovate, 15-17 mm×10 mm, spur subulate, 18-24 mm long, straight or slightly upwards curved, at base 3-4 mm across, lateral sepals subelliptic, 15-18 mm×7-8 mm, lower sepals similar to lateral ones, but larger, 16-20 mm×7-10 mm. Petals 2, blue, glabrous, apex entire. Staminodes 2; limbs blue, orbicular-obovate, 6 mm×4 mm, 2-lobed, adaxially above base densely and shortly yellow-barbate; claws white, 5 mm long, base not appendiculate. Stamens numerous, glabrous; filaments 4-6 mm long, below linear, above filiform; anthers dark-blue, oblong, 0.8-1 mm×0.3-0.5 mm. Carpels 3; ovaries 4 mm long, densely pubescent; styles 2-2.2 mm long, glabrous.

西藏(Xizang)：錯(cuò)那縣，自麻瑪鄉(xiāng)至縣城途中(Cuona Xian,on the way from Mama Xiang to the city of Cuona Xian)，alt. 3 449-4 414 m，山坡，草本植物，花藍(lán)堇色(on slopes, herbs, fls. blue-violet)，2017-09-12，PE西藏考察隊(duì)(PE Xizang Exped.)6429(02293619)(holotype,PE)。

本種與毛莖翠雀花(D.hirticauleFranch.)近緣，與后者的區(qū)別在于本種的莖無(wú)毛或有少數(shù)柔毛，總狀花序具2～4花，花梗在頂端密被短柔毛，退化雄蕊的瓣片圓倒卵形，子房密被柔毛。在毛莖翠雀花，莖下部被長(zhǎng)硬毛，總狀花序具5～10花，花梗無(wú)毛，退化雄蕊的瓣片倒卵形，子房無(wú)毛或近無(wú)毛。(王文采，1979；Wang & Warnock, 2001)

圖3 A～E.扎囊翠雀花：A.莖基部，具一枯萎的最下部莖生葉；B.總狀花序；C.退化雄蕊；D.雄蕊；E.雌蕊群 F～J.寬片翠雀花：F.基生葉；G.總狀花序；H.退化雄蕊；I.雄蕊；J.雌蕊群Fig.3 A-E.Delphinium zhanangense: A.Basal part of stem,with a withered lowermost cauline leaf; B.Raceme; C.Staminode; D.Stamen; E.Gynoecium(from holotype) F-J.D.latilimbum: F.Basal leaf; G.Raceme; H.Staminode; I.Stamen; J.Gynoecium(from holotype)

圖4 狹裂翠雀花 A.開(kāi)花莖；B.退化雄蕊；C.雄蕊；D.雌蕊群；E.基生葉Fig.4 Delphinium lagarolobum A.Flowering stem; B.Staminode; C.Stamen; D.Gynoecium(from holotype); E.Basal leaf(from isotype)

狹裂翠雀花 圖4

DelphiniumlagarolobumW.T.Wang, sp. nov. Fig.4.

(Sect.DelphinastrumDC. )

Ob folia 3-secta dissecta, eorum lobos ultimos linearibus, inflorescentias corymbosas paucifloras, calcar sepalinum subulatum seaplis longius, ovarium pilis densis tectum species nova haec est similisD.caeruleoJacq. ex Camb., a quo foliis semel vel bis divisis, lobis ultimis angustioribus 1-2 mm latis, staminodiorum limbis oblongo-ovatis apice 2-lobulatis, carpellis tribus praeclare differt. InD.caeruleo, folia ter vel quarter divisa, eorum lobi ultimi latiores 1.5-2.5(-4) mm lati, staminodiorum limbi late obovati vel suborbiculares apice integri vel emarginati, et carpella qinque sunt.Perennial herbs. Stems ca. 30 cm tall, with petioles retrorsely appressed-puberulous, 1-2-branched. Basal leaf solitary, long petiolate; blade papery, deltoid in outline, ca. 3 cm×5 cm, at base subtruncate, adaxially glabrous, abaxially on nerves sparsely pubescent, 3-sect and 1-2 times divided, central segment narrow-linear, 3 cm×0.1 cm, near the middle with a small lobule, lateral segments ca. 2.5 cm long, unequally 2-parted to near base, ultimate lobes narrow-linear, upper lobe larger, 2.4 mm×0.1 mm, lower lobe 1.5 cm×0.1 cm; petiole 10.8 cm long. Lower cauline leaves similar to basal one, upper ones with central segment and 2 lateral segments all narrow-linear, 1-2 mm broad, undivided. Corymbs terminal, 2-4.5 cm long, 2-5-flowered; bracts petiolate or sessile, 3-sect, with 3 narrow-linear segments all undivided; pedicels 2-4 cm long, densely puberulous, near or above the middle 2-bracteolate; bracteoles linear, 3-7 mm×0.3-1 mm, puberulous. Flower: Sepals blue, inside glabrous, outside yellowish-puberulous, upper sepal broad-ovate, 14 mm×9 mm, spur subulate, ca. 16 mm long, slightly decurved, at base 4 mm across, lateral sepals obliquely elliptic, 14 mm×8 mm, lower sepals narrow-elliptic, 13 mm×5 mm. Petals 2, blue, glabrous, apex entire. Staminodes 2; limbs blue, oblong-ovate, 7 mm×3.5 mm, at apex 2-lobulate, adaxially below lobules yellow-barbate, above eiliate; claws white, 4 mm long, glabrous, at base 1-appendiculate. Stamens numerous, glabrous; filaments 4-5 mm long, narrow-linear, near apex narrowed; anthers dark-purple, oblong, 1 mm×0.6-0.7 mm. Carpels 3; ovaries 5 mm long, densely white-pubescent; styles blue, 1 mm long.

西藏(Xizang)：隆子縣，玉麥鄉(xiāng)，玉麥雄曲山谷(Lhünzê Xian,Yumai Xiang,Yumaixiongqu Valley)，alt. 2 935 m，山谷林邊，草本植物，花藍(lán)色(forest margin in valley,herbs with blue flowers)，2017-09-06，PE西藏考察隊(duì)(PE Xizang Exped.)5977(02293617)(holotype,PE)。

本種的葉3全裂并細(xì)裂，末回裂片條形，花序傘房狀，具少數(shù)花，萼距鉆形，比萼片長(zhǎng)，子房密被柔毛，因此與也具上述特征的藍(lán)翠雀花(D.caeruleum)相似，與后者的區(qū)別在于本種的葉1～2回分裂，末回裂片較狹窄，寬1～2 mm，退化雄蕊的瓣片長(zhǎng)圓狀卵形，頂端2微裂，心皮3枚。在藍(lán)翠雀花，葉3～4回分裂，末回裂較寬，寬1.5～2.5(～4) mm，退化雄蕊的瓣片寬倒卵形或近圓形，頂端全緣或微凹，心皮5枚。(Hook. f. & Thoms., 1872; 王文采, 1979; Rau, 1993; Wang & Warnock,2001)

致謝蒙馬欣堂、班勤二先生提供珍貴植物標(biāo)本和彩色照片，孫英寶先生為本文繪圖，又蒙鐘小紅女士打印本文稿件，作者在此表示衷心感謝。